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Contributors All authors were involved in defining the content and scope of the paper. TG and SD wrote the first draft of the manuscript. BB collected and interpreted the field and observation data reported. PS prepared the numerical data in its present format and contributed to refinements of the ideas. All authors contributed to amendments of the manuscript and approved the final version.
Funding Princess Alexandra Hospital Foundation Grant and Australian Research Council (DP0880920 and DP140101821).
Competing interests None declared.
Ethics approval Human Research and Ethics Committees of The Princess Alexandra Hospital (HREC/13/QHC/357 and HREC/13/QHC/361).
Provenance and peer review Not commissioned; externally peer reviewed.
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